Chapter 13: The spinal cord, spinal nerves, and somatic reflexes
denticulate ligaments
extend through the arachnoid to the dura, anchoring the cord and limiting side-to-side movements
Tendon reflex
functions when some parts of a muscle contract more than others to spread the workload more evenly over the entire muscle
dorsal root (posterior)
gray matter that carries sensory nerve fibers, which ender the posterior horn of the cord and sometimes synapse with an interneuron there
ventral (anterior) root
gray matter that contain the large somas of the somatic motor neurons. Axons from here exit by way of the anterior root of the spinal nerve and lead to the skeletal muscles
gray commissure
gray matter that the right and left sides of the central core are connected by this
central canal
gray matter this is collapsed in most areas of the adult spinal cord, but in some places (and young children) it remains open, lined with ependymal cells and filled with CSF
white mater
has a bright, pearly white appearance due to an abundance of myelin
Gray matter
has relatively dull color because it contains little myelin
tendon reflex
inhibits alpha motor nurons to the muscle so that it does not contract as strongly
reflexes
play a vital roll in posture, motor coordination, and protective responses to pain or injury
neural integration
pools of spinal neurons receive input from multiple sources, integrate the information, and execute and appropriate output
somatosensory
meaning the nerve carries sensory signals from bones, joints, muscles and the skin
conus medullaris
medullary cone aka
locomotion
motor neurons in the brain initiate walking and determine its speed, distance, and direction
epineurium
protects the nerve from stretching and injury.
reflexes
quick, involuntary, stereotyped reactions of glands or muscles to stimulation
paraplegia
paralysis of both lower limbs resulting from spinal cord lesions at levels T1 to L1
hemiplegia
paralysis of one side of the body, usually resulting from a stroke or other brain lesion
somatic reflexes
reflexes of the skeletal muscle (also called spinal reflexes)
tendon reflex
serves to moderate muscle contraction before it tears the tendon or pulls it loose from the muscle or bone
epineurium
several fascicles are bundled together and wrapped in an ________ to compose the nerve as a whole
central pattern generators
simple repetitive muscle contractions over and over are coordinated by groups of these neurons in the spinal cord
muscle spindles
somatic reflexes involve stretch receptors called _______ ______ embedded in the muscles
subarachnoid space
space between arachnoid membrane and pia mater and is filled with CSF
myotatic
stretch reflex aka
posterior ramus
the _____ _______ innervates the muscles and joints in posterior region of the spin and the skin of the back.
coccygeal ligament
the ______ _____ is formed as the terminal filum and dura mater fuse.
flexor
the _______ reflex employs an ipsilateral reflex arc
spinal nerve proper
the anterior and posterior roots merge, leave the dural sheath and form:
crossed extension reflex
the contraction of extensor muscles in the limb opposite frm the one that is withdrawn. It extends that limb and enables you to keep balanced
extrafusal fibers
the fibers that make up the rest of the muscle and do its work (not intrafusal)
terminal filum
the fibrous strand that continues beyond the medullary cone within the lumbar cistern
intrafusal fibers
the modified muscle fibers within a spindle (at the ends of a muscle, near the tendon)
proprioception
the non visual sense of the position and movements of the body
quadriplegia
the paralysis of all four limbs resulting from lesions above level C5
flexor reflex
the quick contraction of flexor muscles resulting in the withdrawal of a limb from an injurious stimulus
gray matter
the site of all neural integration in the spinal cord
epidural space
the space between the sheath and vertebral bones which is occupied by blood vessels, adipose tissue, and loose connective tissue
conduction
the spinal cords contain bundles of nerve fibers that conduct information up and down the cord, connecting different levels of the trunk with each other and with the brain
meninges
the three fibrous membranes that encloses the brain and spinal cord
spinal nerves
there are 31 _____ ______
5
there are ______ sacral spinal nerves
tendon organs
these are proprioceptors located in a tendon near its junction with a muscle
lower motor neuron
these lead an axon the rest of the way to the muscle or other target organ
corticospinal tracts
these tracts were once called pyramidal tracts
muscle spindles
they inform the brain of muscle length and body movements, which enables the brain to send motor commands back to the muscles that control muscle tone, posture, coordinated movements, and corrective reflexes (balance)
lateral horn
this is present on each side of the gray matter in the spinal cord
lumbar cistern
this is where spinal taps take place
anterolateral system
this system passes up the anterior and lateral columns of the spinal cord
crude touch
touch whose location can only vaguely be identified
dura mater
tough, collagenous membrane that is thick and forms a loose fitting sleeve called the dural sheath around the spinal cord
columns
tracts are arranged in __________ or faniculi
stretch reflex
when a muscle is suddenly stretched and it "fights back"--contracts, increases tone, and feels stiffer than an unstretched muscle
ipsilateral
when a tract does not decussate, its origin and destination are on the same size of the body
monosynaptic reflex arcs
when fibers synapse directly with the alpha motor neurons that return to the muscle, this forms:
spinoreticular tract
Ascending tracts: These tract's third order neurons continue from the pons to the thalamus and fourth-order neurons complete the path to the cerebral cortex
spinothalamic tract
Ascending tracts: Third-order neurons in this tract lead to the cerebral cortex from the thalamus, and arrive in the cerebral hemisphere contralateral to their point of origin
medial lemniscus
Ascending tracts: a tract of nerve fibers that leads the rest of the way up the brainstem to the thalamus
gracile fasciculus
Ascending tracts: an ascending tract that carries signals from the midthoracic and lower parts of the body
spinoreticular tract
Ascending tracts: carries pain signals resulting from tissue injury
cuneate fasciculus
Ascending tracts: carries sensory signals from T6 and up (from the upper limbs and chest)
spinothalamic tract
Ascending tracts: carries signals for pain, temperature, pressure, tickle, itch, and light or crude tough
gracile fasciculus
Ascending tracts: first order nerves that travel up the ipsilateral side of the spinal cord and terminate at the gracile nucleus
spinothalamic tract
Ascending tracts: in this tract, first-order neurons end in the posterior horn of the spinal cord near the point of entry, where they synapse with second-order neurons
cuneate fasciculus
Ascending tracts: occupies the lateral potion of the posterior column and forces the gracile fasciculus medially
Posterior and anterior spinocerebellar tracts
Ascending tracts: their first order neurons originate in muscles and tendons and end in the posterior horn
Posterior and anterior spinocerebellar tracts
Ascending tracts: their second order neurons send their fibers up the spinocerebellar tracts and end in the cerebellum
gracile fasciculus
Ascending tracts: these fibers carry signals for vibration, visceral pain, deep and discriminative touch, and proprioception from lower limbs and trunk
spinothalamic tract
Ascending tracts: these form the anterolateral system
gracile nucleus
Ascending tracts: where gracile fasciculi terminate in the medulla oblongata of the brainstem
T6
At ____ the gracile fasciculus and cuneate fasciculus are joined
corticospinal tracts
Descending tract that carries motor signals from the cerebral cortex for precise, finely coordinated limb movements
tectospinal tract
Descending tract: begins in a midbrain region called the tectum and crosses to the contralateral side of the midbrain
Tectospinal tract
Descending tract: involved in reflex turning of the head, especially in response to sights and sounds
tectospinal tract
Descending tract: it descends through the brainstem to the upper spinal cord on that side, going only as far as the neck.
lateral corticospinal tract
Descending tract: most corticospinal fibers decussate in the lower medulla and for this tract on the contralateral side of the spinal cord
axon
The term "nerve fiber" refers to the ______ of a neuron.
5
There are ____ nerve plexuses.
31
There are ____ pairs of spinal nerves
12
There are ____ thoracic spinal nerves
8
There are _____ cervical spinal nerves
1
There are _____ coccygeal spinal nerve
5
There are _____ lumbar spinal nerves
upper motor neurons
These begin with a soma in the cerebral cortex or brainstem and has an axon that terminates on a lower motor neuron
stretch reflex
This reflex helps to maintain equilibrium and posture
polysynaptic reflex arc
a pathway in which signals travel over many synapses on thier way back to a muscle
reciprocal inhibition
a reflex that prevents muscles from working against each other by inhibiting antagonists
tendon reflex
a response to excessive tension on the tendon
dermatome
a specific area of the skin that sends sensory input to a spinal nerve
endoneurium
a thin sleeve of connective tissue that surrounds nerve fibers
brainstem
a vertical stalk that supports the large cerebellum at the rear of the head and the two cerebral hemispheres that dominate the brain
arachnoid membrane
adhering to the inside of the dura
lumbar puncture
aka spinal tap; a needle is inserted between vertebrae at level L3/L4 or L4/L5, where there is no risk for accidental injury to the spinal cord
coccygeal ligament
anchors the spinal cord and meninges to vertebra
epidural space
anesthetics are sometimes introduced to this space to block pain signals during childbirth or surgery
cauda equina
arising from the lumbar enlargement and medullary cone is a bundle of nerve roots that occupy the ventral canal
Posterior and anterior spinocerebellar tracts
both of these tracts provide the cerebellum with feedback needed to coordinate muscle action
faniculi
columns or _________ of tracts
white matter
composed of tracts that carry signals form one level of the CNS to another
arachnoid mater
consists of a simple squamous epithelium, the arachnoid membrane, and a loose mesh of collagenous and elastic fibers
tendon organ
consists of an encapsulated bundle of small, loose collagen fibers and one or more nerve fibers that penetrate the capsule and end in flattened leaflike processes between the collagen fibers
reflex arc
A somatic reflex employs a _____ _____, which signals travel along the following pathway
integrating center
(reflex arc) a point of synaptic contact between neurons in the gray matter of the cord or brainstem
afferent nerve fibers
(reflex arc) carry information from these receptors to the posterior horn of the spinal cord or to the brainstem
efferent nerve fibers
(reflex arc) carry motor impulses to the muscles and effectors
somatic receptors
(reflex arc) in the skin, muscles and tendons
effectors
(reflex arc) the muscles that carry out a response
reflexes
1. Require stimulation 2. They are quick 3. They are involuntary 4. They are stereotyped (response is very predictable)
gamma
A ________ motor neuron of the spinal cord innervates each end of an intrafusal fiber, preventing it from going slack
posterior and anterior spinocerebellar tracts
Ascending tracts that travel through the lateral column and carry proprioceptive signals from the limbs and trunk to the cerebellum
cuneate nucleus
Ascending tracts: Cuneate fasciculus fibers end here on the ipsilateral side of the medulla oblongata
spinoreticular tract
Ascending tracts: First-Order neurons in this tract immediately synapse with second-order neurons after they ender the posterior horn
medial lemniscus
Ascending tracts: In the medulla, second-order fibers from the gracile and cuneate fasciculi decussate and form this
spinoreticular tract
Ascending tracts: These decussate to the opposite anterolateral system, ascend the cord, and end in a loosely organized core of grey matter (reticular formation) in the medulla and pons
lateral
Descending tract: the ______ vestibulospinal tract passes down the anterior column of the spinal cord and facilitates neurons that control extensor muscles of the limbs, thus inducing the limbs to stiffen and straighten
medial
Descending tract: the _______ vestibulospinal tract splits into ipsilateral and contralateral fibers that descend through the anterior column of both sides of the cord and terminate in the neck. Plays a role in the control of head position
anterior cotricospinal tract
Descending tract: the few corticospinal fibers that do not decussate and remain uncrossed. These are on the ipsialteral side
lateral and medial vestibulospinal tracts
Descending tract: these begin in the brain stem vestibular nuclei, which receive signals for balance from the inner ear
lateral and medial reticulospinal tract
Descending tract: these contain descending analgesic pathways that reduce the transmission of pain signals to the brain
lateral and medial reticulospinal tracts
Descending tract: these control muscles of the upper and lower limbs, especially to maintain posture and balance
later and medial reticulospinal tracts
Descending tract: this originates in the reticular formation of the brainstem
corticospinal tract
Descending tract: this tract gets smaller as it descends and usually disappears by the mid thoracic level
corticospinal tracts
Descending tracts: most of these fibers decussate in the lower medulla and form the lateral corticospinal tract on the contralateral side of the spinal cord
corticospinal tracts
Descending tracts: these fibers form pyramids on the anterior surface of the medulla oblongata
fasciculi
Each column of white matter consists of subdivisions called tracts or ____________
roots; rami
Each spinal nerve branches on both ends into anterior and posterior _______ approaching the spinal cord and anterior and posterior _____ leading away from the vertebral column.
posterior; anterior
Fibers of the _______ tract travel up the ipsilateral side of the spinal cord, and _______ tract fibers cross over and travel up the contralateral side, but then cross back in the brain stem to enter the ipsilateral side of the cerebellum.
decussate
First order neurons synapse with second-order neurons, which ________ and form the contralateral ascending spinothalamic tract. These fibers lead all the way to the thalamus
nerve fibers
First-order neuron, second-order neuron, third order neurons can also be called (first, second, or third) ________ ________
medullary cone
Inferior to the lumbar enlargement is the cord that tapers to a point called the:
brachial
The _____ plexus is a nerve plexus near the shoulder.
meningeal branch
The ______ _____ reenters the vertebral canal and innervates the meninges, vertebrae, and spinal ligaments with sensory and motor fibers
denticulate ligaments
The ______ _______ ligaments are extensions of the pia mater that extend through the arachnoid to the dura mater to anchor the spinal cord.
coccygeal
The ______ plexus is a nerve plexus adjacent to the lower sacrum and coccyx.
cervical
The ______ plexus is a nerve plexus in the neck.
Intercostal nerve
The _______ _____ innervates the internal and external obliques and the transverse abdominal muscles
lumbar
The _______ plexus is a nerve plexus in the lower back
lumbar
The ________ enlargement fives rise to the pelvic region and lower limbs
Sacral
The ________ plexus is a nerve plexus immediately inferior to the lumbar plexus.
cervical
The _________ enlargement gives rise to nerves of the upper limbs.
intercostal nerve
The anterior ramus forms the _______ _____ in the thoracic region, which travels along the inferior margin of a rib and innervates the skin and intercostal muscles (breathing).
contralateral
The crossed extension reflex employs a _______________ reflex arc
anterior ramus
The larger _______ ______ innervates the anterior and lateral skin and muscles of the trunk, and gives rise to nerves of the limbs.
2
The number of nerves in a descending pathway
Foramen Magnum
What structure marks the beginning of the spinal cord?
general
______ fibers innervate widespread organs such as muscles, skin, glands, viscera, and blood vessles
secondary afferent fibers
_______ ______ _______ are sensory nerve fibers in intrafusal fibers in muscle that monitor the length only, not the rate of change
primary afferent fibers
_______ _______ _______ are sensory nerve fibers in intrafusal fibers in muscles that monitor muscle length and how rapidly it changes, which responds to sudden body movements
special
_______ fibers innervate more localized organs in the head, including the eyes, ears, olfactory, and taste receptors, and muscles of chewing, swallowing, and facial expression
alpha
_________ motor neurons supply the extrafusal muscle fibers
ipsilateral
_________ reflex arc: the sensory input and motor output are on the same side of the spinal cord
somatosensory
_________ sensory signals are for touch, heat, cold stretch, pressure, pain, and other sensations.
ascending
_________ tracts carry sensory information up the cord
descending
___________ tracts conduct motor impulses
intersegmental
____________ reflex arc: the input and output occur at different levels of the spinal cord
muscle spindle
a ______ ______ is usually seven or eight small, modified muscle fibers enclosed in an elongated fibrous capsule
nerve
a chordlike organ composed of numerous nerve fibers (axons) bound together by connective tissue
ganglion
a cluster of neurosomas outside the CNS and nerve fibers
spinal cord
a cylinder of nervous tissue that arises from the brainstem at the foramen magnum of the skull
pia mater
a delicate, transparent membrane that closely follows the contours of the spinal cord
ganglion
a knot in a thread of nerves
dural sheath
a loose-fitting sleeve around the spinal cord
tracts
bundles of axons in white matter
descending tracts
carry motor signals down the brainstem and spinal cord
ascending tracts
carry sensory signals up the spinal cord
perineurium
each fascicle is wrapped in a sheath called:
three
how many neurons does it take for a somatic sensation's nerve signal to complete the sensory pathway?
third-order neuron
in an ascending tract, this carries the signal the rest of the way to the cerebral cortex
second-order neuron
in an ascending tract, this continues as far as a gateway called the thalamus at the upper end of the brainstem
first-order neuron
in an ascending tract, this detects a stimulus and transmits a signal to the spinal cord or brainstem
rootlets
in each segment of the cord, six to eight nerve ______ emerge from the anterior surface and converge to form the ventral root of the spinal never
fascicles
in most nerves, the fibers are gathered in bundles called ________
lumbar cistern
inferior to the medullary cone, the subarachnoid space is called the ________ ________ and is occupied by the cauda equine and CSF
perineurium
is composed of up to 20 layers of overlapping, squamous, epithelium-like cells
gray matter
it contains the somas, dendrites, and proximal parts of the axons and neurons
contralateral
when the origin and destination of a tract are on opposite sides of the body
decussation
when tracts cross over from the left side of the body to the right or vice versa